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In the passage "Ghisletta's research used samples from the Berlin Aging Study, which st...

In the passage "Ghisletta's research used samples from the Berlin Aging Study, which started collecting data shortly before the Berlin Wall came down in 1989” (sixth sentence of text 10A2-I), the relative pronoun “which”


A

could be correctly replaced with the relative pronoun “that” without any change in meaning or punctuation.


B

introduces a restrictive clause, meaning the information is essential to identify the study.


C

refers back to the entire preceding clause: "Ghisletta's research used samples from the Berlin Aging Study”.


D

functions as the subject of the verb “started” and refers to the antecedent “the Berlin Aging Study”.


E

functions as the object of the verb “started” within the relative clause.